II. 1. Family Scopuridae

Genus Scopura Uéno, 1929

• Scopura Uéno, 1929, p. 124. Type species: Scopura longa Uéno, by monotypy. Uchida & Maruyama, 1987.

This genus consists of 5 species from Japan (4) and Korea (1). Uchida and Maruyama (1980) provided the diagnoses of the representatives.

1. Scopura bihamulata Uchida, 1987
   • Scopura bihamulata Uchida in Uchida & Maruyama, 1987, p. 704, figs, 6, 12, 18, 22, 28, 34, 40, 49, 52, 56, 59, 64 & 69. Holotype male (NSMT) from Asari-toge, Otaru-shi, Hokkaido.

   
   Distribution: southern Hokkaido.
   Japanese name: FUTAKAGI-TOWADA-KAWAGERA.

    

2. Scopura quadrihamulata Uchida, 1987
   • Scopura quadrihamulata Uchida & Maruyama, 1987, p. 706, figs, 7, 13, 19, 23, 29, 35, 41, 50, 53, 60-1, 65 & 69. Holotype male (NSMT) from Niikappu-ko, Hokkaido.

   
   Distribution: central Hokkaido.
   Japanese name: YOTSUKAGI-TOWADA-KAWAGERA.

    

3. Scopura longa Uéno, 1941
   • Scopura longa Uéno, 1929, p. 125. Holotype a nymph, Wainai, Towada-ko, Akita. Uéno, 1931; Uchida & Maruyama, 1987, p. 702, figs. 1, 5, 10-1, 16-7, 21, 26-7, 32-3, 38-9, 43-4, 46-8, 51, 55, 58, 63, 69.
   • Scopura prolifera Kawai, 1974, p. 275. Holotype male (NSMT), Myoken-zan, Sado Is., Japan. Zwick, 1980.

   [Additainal references: Kohno, 1936, 1937, 1941, 1951a-c, 1953, 1957.]
   
   Distribution: northen Honshu and Sado Island.
   Japanese name: TOWADA-KAWAGERA.

    

4. Scopura montana Maruyama, 1987
   • Scopura longa : Uéno, 1931, p. 40, fig. 3. In part (adult). Kawai, 1967, p. 4, 178, figs. 105-8; Zwick, 1973, p. 167, fig. 57; Kawai, 1974, p. 275, fig. 2 d-f. [Additinal references: Uéno, 1935, 1938, Kobayashi, 1981; Uchida, 1981, 1984; Zwick, 1980; Kawai & Isobe, 1983).
   • Scopura montana Maruyama in Uchida & Maruyama, 1987, p. 701, fig. 2, 4, 8-9, 14-5, 20, 24-5, 30-1, 36-7, 42, 45, 54, 57, 62, 68 & 69. Holotype, male (NSMT) Mito-sawa, Hinoharamura, Tokyo, Japan. Uchida, 1989; Ida, 1994; Kishimoto, 1997.

   
   [Additainal references: Komatsu, 1955, 1956, 1957, 1958, 1959; Komatsu & Takahashi, 1952.]
   
   Distribution: central Honshu.
   Japanese name: MINE-TOWADA-KAWAGERA.

REFERENCES

 

• IDA, M. 1994. The life cycle of Scopura montana (Plecoptera: Scopuridae). Japanese Journal of Limnology, 55: 23-25.
• KAWAI, T. 1967. Plecoptera (Insecta). Biogeographical Society of Japan, Tokyo.
• KAWAI, T. 1974. The second species of the genus Scopura. Bulletin of the National Science Museum, Tokyo, 17: 285-281.
• KAWAI, T. & ISOBE, Y. 1983. Notes on the egg of Scopura longa Uéno (Plecoptera). Annales de Limnologie, 20: 57-58.
• KOBAYASHI, M. 1981. New locarity of Scopura longa Uéno. Natural History Reports of Kanagawa, Yokohama, 2: 45-46.
• KISHIMOTO, T. 1997. Egg envelopes of a stonefly Scopura montana Maruyama (Plecoptera: Scopuridae). Proc. Arthropod. Embryol. Soc. Jpn., 32: 35-37.
• KOHNO, M. 1936. Description of the nymph of Peltoperla sp., and a record of a new locality of Scopura longa Uéno. Kontyû, 10: 103-108.
• KOHNO, M. 1937. Notes on the Perlodes yarizawana Uéno and Scopura longa Uéno. Mushi, 9: 116-119.
• KOHNO, M. 1941. The breeding temprature and the food of Scopura longa Uéno (larva). Mushi, 14: 33.
• KOHNO, M. 1951. New locality of Scopura longa Uéno. Shin Kontyû, 4: 10-11.
• KOHNO, M. 1951. Notes on Scopura longa Uéno of Aizu, Fukushima Pref. (1). Collecting and Breeding, 13: 49-54.
• KOHNO, M. 1951. Notes on Scopura longa Uéno of Aizu, Fukushima Pref. (2). Collecting and Breeding, 13: 128-130.
• KOHNO, M. 1953. New locality of Scopura longa Uéno, recorded by Mr. Ikeda. Shin Kontyû, 6: 51.
• KOHNO, M. 1957. A new locality of Scopura longa Uéno. Shin Kontyû, 10: 43.
• KOMATSU, T. 1955. Studies on the emergence of Scopura longa Uéno. Shin Kontyû, 4: 1-8.
• KOMATSU, T. 1956. On the imago, egg and first instar of Scopura longa Uéno. Shin Kontyû, 5: 13-21.
• KOMATSU, T. 1957. Notes on the Scopura longa Uéno. Shinano Seibutsu, : 5-18.
• KOMATSU, T. 1958. Some contributions to the knowledge of Scopura longa Uéno. Animals of North Shinano (Kitashinano no Seibutu), 1: 1-15.
• KOMATSU, T. 1959. On the thermal reaction of the adult Scopura longa Uéno (Plecoptera). Shin Kontyû, 8: 23-28.
• KOMATSU, T. & TAKAHASHI, K. 1952. Distribution of Scopura longa Uéno in northern Shinano. Collecting and Breeding, 14: 357-359.
• MIYAKO, T. 1956. Allometric growth of the nymphs of Scopura longa Uéno. New Entomologist, Ueda, 5: 27-29.
• OCHIAI, T. 1953. Localities of Scopura longa in Nagano Prefecture. Shin Kontyû, 6: 39-40.
• UCHIDA, S. 1981. Benthic animals of mountain streams in the Oi gawa genrybu wilderness area in the southern Japan. In Conservation Reports of the Oi gawa Genrybu Wilderness Area in the Southern Japanese Alps, Central Japan.
• UCHIDA, S. 1984. Distribution of the large stoneflies in Tanzawa mountains, Natural History Reports of Kanagawa, Yokohama, 5: 17-25.
• UCHIDA, S. 1989. Distribution of Plecoptera in the Tama-Gawa River system, central Japan. In Campbell, I. C. (ed.), Mayflies and stoneflies: Life histories and biology. Ser. Entomol. vol. 44 : 181-188. Kluwer Academic, Dordrecht (Netherlands).
• UCHIDA, S. & MARUYAMA, H. 1987. What is Scopura longa Uéno, 1929 (Insecta, Plecoptera)? A revision of the genus. Zoological Science, 4: 699-709.
• UÉNO, M. 1929. Studies on the Stoneflies of Japan. Memoirs of the College of Science, Kyoto Imperial University, Series B, 4: 97-115.
• UÉNO, M. 1931. Einige neue Ephemeropteren und Plecopteren aus Mittl Japan. Annotationes Zoologicae Japonses, 16.
• UÉNO, M. 1935. Plecoptera. In "Aquatic animals of the Azusa-gawa river system and Kamikochi." Iwanami, Tokyo, pp. 30-51.
• UÉNO, M. 1938. Scopuridae, an aberrant Family of the Order Plecoptera. Insecta Matsumurana, 7: 154-159.
• ZWICK, P. 1973. Insecta: Plecoptera, Phylogenetisches System und Katalog. Walter de Gruyter, Berlin.
• ZWICK, P. 1980. Plecoptera (Steinfliegen). Walter de Gruyter, Berline.